Hybrid origin
Northeastern United States, Mid-Atlantic United States, Southeastern United States, Central United States
boissieri , cupanii , hungaricum , kesselringii
Ardis, Arnold, Aureomarginatum
Temperate Regions
Bluffs, low mountains, Moist Soils, Woodlands
21,22
1a, 1b, 2a, 2b, 3a, 3b, 4, 5, 6, 7, 8, 9, 10, 11, 12, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23